Undergraduate Education: BA, Molecular Biology, San Jose State University, 1979
Graduate Education: Ph.D., Biochemistry and Molecular Biology, University of California, Santa Barbara, 1986
Post-doctoral education: Neuroscience Research Institute, University of California, Santa Barbara
Previous Faculty Positions: Asst. Prof, Assoc. Prof., Professor, Department of Ophthalmology, Medical College of Wisconsin (1991 – 2008)
Memberships: International Color Vision Society (ICVS), Association for Research in Vision and Ophthalmology (ARVO)